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Cardellini received her first big break role in 1996 when she landed a starring role as Sarah on ABC's Saturday morning live-action children's series, Bone Chillers.Following this, she made guest appearances on prime-time programs such as Step by Step, Clueless, 3rd Rock from the Sun, and Boy Meets World as Lauren, a girl that came between the show's star couple.She joined the cast of the hospital drama ER in 2003 as Samantha Taggart, a free-spirited nurse.Cardellini acted for six seasons on ER, until the series finale, "And in the End...", and with the cast won the TV Land ensemble Icon Award in 2009.Cardellini starred in the live-action adaptation of Scooby-Doo in 2002, in which she played the cartoon character Velma Dinkley.
Mystery Incorporated, Wendy Corduroy in Gravity Falls, and Megan in Sanjay and Craig.
She appeared with cast members and producers of Freaks and Geeks and Undeclared at the Paley Center for Media Paley Fest on March 12, 2011.
From June 15, 2012 to February 15, 2016, she has been the voice of Wendy on the Disney Channel show Gravity Falls.
Cardellini voice acts periodically on the podcast and live show The Thrilling Adventure Hour, on the Sparks Nevada: Marshal on Mars segments.
In 2013, she was chosen to play Sylvia Rosen, a love interest of Don Draper on Mad Men.